Manual of California Native Plants
Easy Companion Planting: The most important thing to know about.
"Let wind or storm, axe or fire, isolate trees of the forests and they are doomed; they are ... unable to adapt themselves to changing conditions...So long as their normal life is not interfered with all is well, but if brought face to face with changed conditions and adversities they have no reserve of adaptability and fall by the wayside." Ernest Wilson (about 1920) (Foley)
Plants grow in many diverse sites throughout the world. They have developed different ways to get nutrition and moisture from each soil and climate. They have to deal with different animals eating them in different areas. This causes many problems when you plant plants that have no protection, no ties, no food and no water strategies for your plant community, soil or climate.
Plants have evolved to different strategies for each plant community, soil type and climate. That is a reason you do not often see only one type of plant in the wild over vast areas. This is not competition in the old sense. The only places 'competition' exists are in your yard, wet disturbed places or weedy areas. (Most yards fit this competitive scenario because we grow plants from many plant communities all mixed together.) In the wild, competition per say does not exist. The real world relies on redundancy, help from friends in the community and a very long-term look at life. The individuals become part of the whole and live and die with it.
Below are the different strategies that forms of life use to survive: (adapted from Grime, 1979)
Realize these are not set in stone but are generalities and tendencies.
Ruderals
Ruderals, "R"
or "r" selection. These are the weeds of the world.
Their whole life is spent trying to make as many seeds as possible
before their short life is spent. Ruderals are eaten by all as energy
is not used for defense but only to produce flowers and seed. Ruderals
need very high nutrition and regular water to stay alive. Most are
not community oriented. (Ruderals are non-mycorrhizal or will become mycorrhizal only if food or water can be
drawn from the mycorrhizal grid. Mycorrhiza means fungus root. 90% of
the plants in the world have a fungal partner that helps them draw
nutrition and moisture from the soil. Mycorrhizas protect plants from
diseases and insects and can tie vast areas together.) Ruderals flower
like mad (trying to produce seeds). Their many flowers and fruits make them the
annual color plants and many vegetables that we plant in our gardens.
Ruderals have a limited ability to grow well in shade. Ruderals are the
first ones into disturbed areas where they use as much water and
nutrition as fast as possible. Ruderals like bare ground and cannot
reseed well on mulched areas. Their growth is not helped with mulch and
Ruderals will sometimes rot off at the stem if they are mulched.
Ruderals live in a bacterial-based soil that can often be wet, soggy
and highly fertile. (If only for as long as it takes to make seed.) Do not mix ruderals with most California native plants.
R simply defined
High nutritional needs
High water use(inefficiently)
Short life
Lots of flowers
Hard to kill or control (as it will seed in days if the conditions are right)
Insects damage ( think of all the herbicides people use in their flower gardens)
Likes soil amendments, hates mulch
Many seedlings
Weedy' growth
Very competitive
Individualistic to
their species, I.e., the plant community is not supported, ruderals try
to collapse community structure (a single weed in the pot with a
mycorrhizal species is normally fatal.)
Some Common Ruderals
Alyssum, Creeping Charlie ,Beets ,Dicentra, Purslane, Dusty Miller, Radish, Bouncing Bet, Four-O'Clock ,Rape ,Broccoli, Garden Cress, Rhubarb,Cabbage, Globe-Amaranth, Sandwort, Candytufts, Ice plant ,Starlings, Sorrel, Carnation, Kale, Spinach, Cauliflower, Kohlrabi ,Stock Chard, Lawns(except Bermuda), Stone-cress, Collards ,Mustard, Sweet William, Nasturtium, Turnips ,New Zealand Spinach, Wallflower, Pinks, Wandering Jew, Poke-weed
Circumventer
Circumventer ,"C"
selection (circumventor or circumventer). These are many of our western
wildflowers, perennials, shrubs, and much of the rest of the worlds
shrubs and trees. These are the plants that shut down during stress
(sometimes for years or decades.) When there is much nutrition or
moisture available Circumventers grow, flower and produce seed, in much
of the west that is our spring or early summer. The rest of the year
Circumventers are dormant. When Circumventers go through this cycle,
they produce much mulch. Cs are many of the secondary fire-followers
that rebuild the mulch on the disturbed site. Circumventers usually are
not bothered much by bugs or animals during their spring flush, but if
you water them during the summer and keep them evergreen expect them to
be unstable and have some bugs. In other areas of the country that have
much summer rain these are most of the shrubs and trees. The trees grow
tall and fast and are largely deciduous. These plants can live a long
time. Circumventers store some energy and do not rely only on yearly
seed production to live from year to year. The perennials can live for
10 to 150 years. Our wildflowers are annual forms of these.
Circumventers will sometimes lie dormant for years before germinating.
(That is a reason why wildflower seed remains viable for years but some
vegetable seeds fail after a couple of years.) Most of the
circumventive perennials, shrubs and trees tolerate or require surface
mulch. Circumventers can handle regular water only when they are used
to it in their wild environment.
Circumventers that have stress-tolerant or ruderal tendencies will sometimes change when they are introduced into another climate, soil or plant community. If the limiting stress is removed, a borderline stress tolerant may be a full stress tolerant, (Ceanothus in many spots) if water was making a plant a circumventer regular water may allow it to become more ruderal (Ca. Poppies in high rainfall areas). A C plant in the mellowest community it lives in it may become a K.
R -C- K
C= Many flowers during their flowering period and usually are not weedy
community oriented(provides support for the fungal grid and animals)
Cs can be very drought tolerant, particularly if associated with its native neighbors or in the shade of its companion trees.
Do not underestimate the 'disappearance' under stress. Some species do not even leave a bump. It seems a miracle when it reappears when the stress is removed. Amphibians fit this, as does the poker player that folds a lot waiting for the perfect hand.
(A poker playing frog?)
Some Common Circumventers
Acacia, Corn, Mock Orange, Tulip Tree ,Alders, Currants, Most deciduous trees ,Viburnums, Ash, Dogwood, fruit trees, Wheat, Asters, Elm ,Nut Trees, Baby Blue eyes, Forsythia, Oleander, Barley, Fuchsia, Onions, Tomatoes, Bermuda grass, Gingko, Penstemons, Berries, Gooseberries, Plane, Buckeye, Hibiscus, Potato Bulbs, most Hydrangea, Privet Lupine Calif. Poppy Iris Chrysanthemum Liquidambar Roses Spiraea Maple Bougainvillea
Stress-tolerant
Stress-tolerant, "S" or "k" selection. In highly stressed environments these are most of the plants that can survive longer than a season. Stress-tolerants grow where the soil is poor, the rainfall is low, the climate harsh, or the sunlight is low or limited. These plants have evolved slow, long-term actions and live long, to very long. The perennial forms of these can live for a hundred or more years. The shrubs and trees can live for a thousand years. Stress-tolerants may make seeds only every 50 years or so, or Stress-tolerants may make only small quantities every year expecting a fire or other mass disturbance to germinate them sometime in a 100-500 year period. Stress-tolerants may carry each leaf for years. Stress-tolerants have many redundant strategies to survive, e.g., crown sprout, seed and sucker to reproduce. Stress-tolerants have thick bark in fire areas and lie low on the ground in arctic areas. Stress-tolerants are mutualistic, (i.e., help each other) and highly mycorrhizal, sharing their strengths with each other within their community, passing nutrients and moisture underground. The stress-tolerant creek species that grow next to creeks have water but may be limited by nutrition (because of high seasonal moisture), while the ones on the hillside are limited by water but not nutrition; by sharing, each is stronger. Stress-tolerant plants cannot handle high moisture and nutrition other than seasonally when their fungal partners are inactive(only for 2 months or so each year). Stress-tolerants love surface mulch. Desert ones like rock, forest ones like chipped wood or leaves(litter). Stress-tolerants often have built-in weed control. That is, Stress-tolerants limit their own growth and the growth of plants nearby to use the nutrients and water available. Stress-tolerants live in a fungal-based soil.
Some common Stress
Tolerants Bay, California Fir Blueberries Incense Cedar Bunch
Grasses(interior
forms) Ceanothus(interior forms) Manzanitas Cranberries
Oaks, evergreen Creosote Cypress Photinia Douglas Fir Pines Eucalyptus
S or k= Long lived
drought tolerant if part of the community
doesn't crave water other than what, when, and how it occurs in its natural community
(plants from the lower desert drown in high winter rains, most of our plants hate summer rainfall or watering.)
usually bug free likes low fertility mulch hates being fertilized hates amended soil hates insecticides or fungicides
The best S or K plant is one most adapted to your site, i.e., native on the site.
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Las Pilitas Nursery |
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Santa Margarita - Escondido |